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Your account number is a number that uniquely identifies your account with a bank, credit card company, etc. It's one-of-a-kind. For example, the account number on a credit card is the big number embossed on the front of your card. On loans, it can usually be found on your account statement.
Don't lose it, share it, or say it on the phone unless you called them. Never send it via unsecured email, either.